While attending a live dubbing of his favorite anime, Aruto has to protect the leading HumaGear actress from A.I.M.S., who have come to arrest her because her manager broke a HumaGear law by basing her on his deceased daughter. When MetsubouJinrai.net attack, Aruto loses his Flying Falcon Progrise Key during the ensuing fight, which results in a new Kamen Rider appearing on the scene to challenge Aruto as Zero-One. Meanwhile, Yua's mysterious associate gives her the Lightning Hornet Progrise Key to assist her in battle.